0001001 - Document - Cover link:presentationLink link:calculationLink link:definitionLink 0002002 - Document - Audit Information link:presentationLink link:calculationLink link:definitionLink 1001003 - Statement - Consolidated Balance Sheets link:presentationLink link:calculationLink link:definitionLink 1002004 - Statement - Consolidated Balance Sheets (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 1003005 - Statement - Consolidated Statements of Operations link:presentationLink link:calculationLink link:definitionLink 1004006 - Statement - Consolidated Statements of Convertible Preferred Stock and Stockholders’ Equity (Deficit) link:presentationLink link:calculationLink link:definitionLink 1005007 - Statement - Consolidated Statements of Convertible Preferred Stock and Stockholders’ Equity (Deficit) (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 1006008 - Statement - Consolidated Statements of Cash Flows link:presentationLink link:calculationLink link:definitionLink 1407401 - Statement - Consolidated Statements of Cash Flows (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 2101101 - Disclosure - Nature of Business and Basis of Presentation link:presentationLink link:calculationLink link:definitionLink 2402402 - Disclosure - Nature of Business and Basis of Presentation (Details) link:presentationLink link:calculationLink link:definitionLink 2103102 - Disclosure - Summary of Significant Accounting Policies link:presentationLink link:calculationLink link:definitionLink 2204201 - Disclosure - Summary of Significant Accounting Policies (Policies) link:presentationLink link:calculationLink link:definitionLink 2305301 - Disclosure - Summary of Significant Accounting Policies (Tables) link:presentationLink link:calculationLink link:definitionLink 2406403 - Disclosure - Summary of Significant Accounting Policies - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 2407404 - Disclosure - Summary of Significant Accounting Policies - Property and Equipment (Details) link:presentationLink link:calculationLink link:definitionLink 2408405 - Disclosure - Summary of Significant Accounting Policies- Antidilutive Securities (Details) link:presentationLink link:calculationLink link:definitionLink 2109103 - Disclosure - Fair Value Measurements link:presentationLink link:calculationLink link:definitionLink 2310302 - Disclosure - Fair Value Measurements (Tables) link:presentationLink link:calculationLink link:definitionLink 2411406 - Disclosure - Fair Value Measurements - Assets and Liabilities Measured at Fair Value (Details) link:presentationLink link:calculationLink link:definitionLink 2412407 - Disclosure - Fair Value Measurements - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 2413408 - Disclosure - Fair Value Measurements - Significant Quantitative Inputs (Details) link:presentationLink link:calculationLink link:definitionLink 2414409 - Disclosure - Fair Value Measurements - Changes in Level 3 Fair Value Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 2115104 - Disclosure - Prepaid Expenses and Other Current Assets link:presentationLink link:calculationLink link:definitionLink 2316303 - Disclosure - Prepaid Expenses and Other Current Assets (Tables) link:presentationLink link:calculationLink link:definitionLink 2417410 - Disclosure - Prepaid Expenses and Other Current Assets (Details) link:presentationLink link:calculationLink link:definitionLink 2118105 - Disclosure - Property and Equipment, Net link:presentationLink link:calculationLink link:definitionLink 2319304 - Disclosure - Property and Equipment, Net (Tables) link:presentationLink link:calculationLink link:definitionLink 2420411 - Disclosure - Property and Equipment, Net - Summary of Property and Equipment, Net (Details) link:presentationLink link:calculationLink link:definitionLink 2421412 - Disclosure - Property and Equipment, Net - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 2122106 - Disclosure - Accrued Expenses and Other Current Liabilities link:presentationLink link:calculationLink link:definitionLink 2323305 - Disclosure - Accrued Expenses and Other Current Liabilities (Tables) link:presentationLink link:calculationLink link:definitionLink 2424413 - Disclosure - Accrued Expenses and Other Current Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 2125107 - Disclosure - License Agreements link:presentationLink link:calculationLink link:definitionLink 2426414 - Disclosure - License Agreements (Details) link:presentationLink link:calculationLink link:definitionLink 2127108 - Disclosure - Convertible Preferred Stock link:presentationLink link:calculationLink link:definitionLink 2428415 - Disclosure - Convertible Preferred Stock - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 2129109 - Disclosure - Common Stock link:presentationLink link:calculationLink link:definitionLink 2330306 - Disclosure - Common Stock (Tables) link:presentationLink link:calculationLink link:definitionLink 2431416 - Disclosure - Common Stock - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 2432417 - Disclosure - Common Stock - Summary of Common Stock Reserved for Future Issuance (Details) link:presentationLink link:calculationLink link:definitionLink 2133110 - Disclosure - Stock-Based Compensation link:presentationLink link:calculationLink link:definitionLink 2334307 - Disclosure - Stock-Based Compensation (Tables) link:presentationLink link:calculationLink link:definitionLink 2435418 - Disclosure - Stock-Based Compensation - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 2436419 - Disclosure - Stock-Based Compensation - Stock Option Valuation (Details) link:presentationLink link:calculationLink link:definitionLink 2437420 - Disclosure - Stock-Based Compensation - Stock Option Activity (Details) link:presentationLink link:calculationLink link:definitionLink 2438421 - Disclosure - Stock-Based Compensation - Restricted Stock Activity (Details) link:presentationLink link:calculationLink link:definitionLink 2439422 - Disclosure - Stock-Based Compensation - Stock-Based Compensation Expense (Details) link:presentationLink link:calculationLink link:definitionLink 2140111 - Disclosure - Income Taxes link:presentationLink link:calculationLink link:definitionLink 2341308 - Disclosure - Income Taxes (Tables) link:presentationLink link:calculationLink link:definitionLink 2442423 - Disclosure - Income Taxes - Schedule of Loss Before Provision For (Benefit From) Income Taxes (Details) link:presentationLink link:calculationLink link:definitionLink 2443424 - Disclosure - Income Taxes - Schedule of Components of Income Tax Expense (Benefit) (Details) link:presentationLink link:calculationLink link:definitionLink 2444425 - Disclosure - Income Taxes - Reconciliation of U.S. Federal Statutory Income Tax Rate to Effective Income Tax Rate (Details) link:presentationLink link:calculationLink link:definitionLink 2445426 - Disclosure - Income Taxes - Schedule of Net Deferred Tax Assets (Details) link:presentationLink link:calculationLink link:definitionLink 2446427 - Disclosure - Income Taxes - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 2147112 - Disclosure - Commitment and Contingencies link:presentationLink link:calculationLink link:definitionLink 2348309 - Disclosure - Commitment and Contingencies (Tables) link:presentationLink link:calculationLink link:definitionLink 2449428 - Disclosure - Commitment and Contingencies - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 2450429 - Disclosure - Commitment and Contingencies - Summary of Lease Costs (Details) link:presentationLink link:calculationLink link:definitionLink 2451430 - Disclosure - Commitment and Contingencies - Maturity of Lease Liability (Details) link:presentationLink link:calculationLink link:definitionLink 2451430 - Disclosure - Commitment and Contingencies - Maturity of Lease Liability (Details) link:presentationLink link:calculationLink link:definitionLink 2452431 - Disclosure - Commitment and Contingencies - Future Payments Under Loan Obligation (Details) link:presentationLink link:calculationLink link:definitionLink 2153113 - Disclosure - Revenue from Contracts with Customers link:presentationLink link:calculationLink link:definitionLink 2454432 - Disclosure - Revenue from Contracts with Customers (Details) link:presentationLink link:calculationLink link:definitionLink